Botanic Gardens Trust, Sydney, Australia

Helen Ramsay

Helen Ramsay MSc, PhD(Syd)

Honorary Research Associate


(02) 9869 1447

Research interests
Bryophytes (mosses)

Mosses, taxonomy, genetics, cytology, bryophytes

Current research projects

  • Checking the identity of names listed as TYPES for the genus Trematodon
  • Identifying and returning loans of Bryaceae from MEL
  • Checking specimens from Utrecht (Prof Ben van Zanten) and returning unrequired ones.
  • Identifying and checking specimens from Bryaceae, Orthotrichaceae and Sematophyllaceae
  • Possible collaborative project for publication

Key publications

  • Brown, E.A., H.P. Ramsay & J. Seur. 1992. Australian hepatic types at the National Herbarium of New South wales (NSW) and the Ray Herbarium in Sydney (SYD). Telopea 5(12): 67-90.
  • Downing, A.J., H.P. Ramsay & W.B. Schofield. 1991. Bryophytes in the vicinity of Jenolan Caves. Cunninghamia 2(3): 371-384.
  • Ramsay, H.P. 1973. Unusual sporocytes in Dicnemoloma pallidum (Hook.) Wijk & Marg. Bryologist 76: 178-182.
  • Ramsay, H.P.& B.G. Briggs 1979. The bryophyte and lichen collections at the National Herbarium of New South Wales. Australian Systematic Society Newsletter 18: 13-14.
  • Ramsay, H.P. & B.G. Briggs. 1979. The bryophyte collections at NSW, Sydney. Taxon 28: 639-640.  
  • Ramsay, H.P  1979. Anisospory and sexual dimorphism in the Musci. In. Clarke, G.C.S.  & J.G. Duckett (eds). Bryophyte Systematics. Syst. Assoc. Special volume 14: 281-326. Academic Press, London.
  • Ramsay, H.P. 1980. Contributions of Rev. W.W. Watts F.L.S. to Australian Botany. Taxon 29(4): 455-469.
  • Ramsay, H.P. 1984.  Census of New South Wales Mosses. Telopea 2(5): 455-533
  • Ramsay, H.P. 1984.  Phytogeography of  the mosses of New South Wales. Telopea 2 (5): 534-548.
  • Ramsay, H.P. & H. Streimann 1984. Mosses and their distribution in the A.C.T. Telopea 2(5): 559-574.
  • Ramsay, H.P. 1984. The mosses of Lord Howe Island. Telopea 2(5): 549-558
  • Ramsay, H.P. & D.H. Vitt. 1984. Distribution, cytotaxonomy and sexuality of Macromitrium in the Pacific. J. Hattori Bot. Lab.  55: 23-33.
  • Ramsay, H.P. 1986. Studies on the moss Holomitrium perichaetale. Hikobia 9(4): 307-314.
  • Ramsay, H.P., H. Streimann, A Ratkowsky, R. Seppelt, & A. Fife. 1986. Australasian alpine bryophytes. In. Barlow, B. (ed) . Flora and fauna of  Alpine Australasia.: Ages and origins. Chap. 20. 301-335. CSIRO Melbourne.
  • Ramsay, H.P., H. Streimann & G. Harden. 1987. Observations on the bryoflora of Australasian rainforests. Acta Bot. Hungarica 35: 605-620.
  • Ramsay, H.P. 1988. Studies on the reproductive biology of some mosses from the Australian wet tropics.  Proc. Ecol. Soc. Aust. 15: 273-279.
  • Ramsay, H.P & J. Seur. 1990. Type specimens of bryophytes in Australian Herbaria. I. P.S. Short (ed.) History of Systematic Botany in Australasia. Australian Systematic Society Inc. Melbourne pp. 247-252.
  • Ramsay, H.P., J. Seur, P.G. Wilson & T. Goodwin. 1990. Register of type specimens of mosses in Australian Herbaria. Part. 1. General Introduction and special collections at NSW: Lord Howe Island and Vanuatu (New Hebrides. Telopea 3(4): 571-592.
  • Ramsay, H.P., A.J. Downing & W.B. Schofield 1991. The bryophytes of Mt Tomah Botanic Garden. Cunninghamia 2(3): 295-304.
  • Ramsay, H.P., R.G. Coveny, E.A.Brown and A.K. Brooks. 1993. Bryophytes of the Royal Botanic Gardens, Government House and the Domain, Sydney. Cunninghamia 3(1): 215-229.
  • Ramsay, H.P.& J. Seur. 1994.  Register of Type Specimens of Mosses in Australian Herbaria. Flora of Australia.  Supplementary Series  No 2. ABRS Canberra
  • Ramsay,  H.P., W.B. Schofield and B.C.Tan  2002.  The genus Taxithelium. (Bryopsida: Sematophyllaceae) in Australia. Austral. Syst. Bot.15: 583-596.
  • Ramsay, H.P., W.B. Schofield, and B.C. Tan. 2002 . The family Sematophyllaceae (Bryopsida) in Australia. Part 1. Introduction, family data, kley to the genera and the genera Wijkia, Acanthorrhynchium, Trismegistia and  Sematophyllum. J. Hattori Bot. Lab. 92: 1-50.
  • Ramsay, H.P., W.B. Schofield and B. C. Tan 2004. The family Sematophyllaceae (Bryopsida) in Australia. Part 2. Acroporium, Clastobryum, Macrohymenium, Meiotheciella, Meiothecium, Papillidiopsis, Radulina, Rhaphidorrhynchium, Trichosteleum,  and Warburgiella. J. Hattori Bot. Lab. 95: 1-69.
  • Spence, J.R. & H.P. Ramsay 1996 Biogeography of the Subfamily Bryoideae (Bryaceae, Musei) in north-eastern Queensland. Australian Systematic Botany 9: 185-192.
  • Spence, J.R. & H.P. Ramsay 1999. Three new species of Rosulabryum (Bryopsida, Bryaceae) from Australia.  Telopea 8(3): 325-335.
  • Spence, J.R. & H.P. Ramsay 1999. Proposal to conserve the name Bryum (Musci,  Bryaceae) with a conserved type. Taxon: 827-828.
  • Spence, J.R. & H.P. Ramsay 1996. New and interesting species of the family Bryaceae (Bryopsida) from Australia. J. Adelaide Bot. Gard. 17: 107-118.
  • Spence, J.R & H.P. Ramsay 2002 The genus Anomoliryum Schimp (Bryopsida, Bryaceae) in Australia. Telopea 9(4): 777-792.
  • Tan, B.C., H.P. Ramsay & W.B. Schofield. 1996. A contribution to Australian Sematophyllaceae (Bryopsida).  Austral. Syst. Bot. 9: 319-327.
  • Tan, B.C., W.B. Schofield & H.P. Ramsay 1998. Miscellanies of Australian Sematophyllaceae with a new genus, Meiotheciella. Nova Hedwigia 67(1-2): 213-223.
  • Vitt, D.H. & Ramsay, H.P.1984.  A cladistic analysis of the moss Macromitrium in Australasia. Aust. Syst. Bot. Soc. 40:25.
  • Vitt, D.H. & H.P. Ramsay 1985. The Macromitrium complex in Australasia (Orthotrichaceae:Bryopsida) Part. I.  Taxonomy and phylogenetic relationships. J. Hattori Bot. Lab. 59: 325-451
  • Vitt, D.H. & H.P. Ramsay 1985  The Macromitrium complex in Australasia (Orthotrichaceae: Bryopsida). Part II. Distribution, Ecology, and Paleogeography. J. Hattori Bot. Lab. 59: 453-468. 

Chromosome studies

  • Ramsay, H.P. 1963  Chromosome numbers in the Proteaceae. Aust. J. Bot. 11: 1-20.
  • Ramsay, H.P.1964. The chromosomes of Dawsonia. Bryologist 67: 153-162.
  • Ramsay, H.P.1966.  Sex chromosomes in Macromitrium. Bryologist 69: 293-311.
  • Ramsay, H.P. 1969. Cytological studies on some mosses from the British Isles. Bot. J. Linn. Soc. 62: 85-121.
  • Ramsay, H.P. 1974. Cytological studies on Australian mosses. Austral. J. Bot.22: 293-348.
  • Ramsay, H.P. & W.B. Schofield. 1981. Observations on the cytology of mosses endemic to Western North America which occur in British Columbia. J. Hattori Bot. Lab. 49: 279-304
  • Ramsay, H.P. 1982. The value of karyotype analysis in the study of mosses. J. Hattori Bot. Lab. 53: 51-71.
  • Ramsay, H.P. 1983. A moss with four chromsomes. Lindbergia 9: 89-92.
  • Ramsay, H.P. 1983. Chromosome numbers in some mosses from western Canada.Can. J. Bot. 61: 2384-2387.
  • Ramsay, H.P. 1983. Cytology of Mosses. In. Schuster R.M. (ed.) New Manual of Bryology. Pp 149-221. Hattori Botanical Laboratory, Nichinan.
  • Ramsay, H.P,. & J. Lewinsky. 1984. Chromosome studies on some Australasian Orthotrichaceae (Musci). 1. Orthotrichum. New Zealand J.Bot. 22: 346-351.
  • Ramsay, H.P. & D.H. Vitt. 1984. Distribution, cytotaxonomy and sexuality of Macromitrium in the Pacific. J. Hattori Bot. Lab. 55: 23-33.
  • Ramsay H.P. 1985. Cytological and sexual characteristics of the moss Dicranoloma. In Crosby, M.R. (ed.). Studies in Systematic Botany dedicated to Lewis E.  Anderson. Monographs in Systematic Botany from the Missouri Botanical Garden .2: 93-110.
  • Ramsay, H.P. & D.H. Vitt. 1986. The Macromitrium complex in Australasia (Orthotrichaceae: Bryopsida) Part III. Cytotaxonomy. J. Hattori Bot. Lab. 61: 1-43.
  • Ramsay, H.P.  1987. Cytological and other studies on the Hypnodendraceae. Memoirs New York Botanical Garden 45: 135-153.
  • Ramsay, Helen P. 1988. Cytological and biosystematic studies on some Australasian mosses. J. Hattori Bot. Lab. 64: 177-186.
  • Ramsay, H.P. 1992. Chromosome studies on some mosses from Lord Howe Island. Nova Hedwigia 54 (1-2).: 37-43.
  • Ramsay, H.P. & J. Lewinsky. 1993. Chromosome studies on some Australasian Orthotrichaceae I. Ulota and Zygodon with additional studies on Orthotrichaum, Schlotheimia and Macromitrium. J. Hattori Bot. Lab. 74: 183-192.
  • Ramsay, H.P., H. Streimann & D.H. vitt. 1995. Cytological studies in mosses from Papua New Guinea 1. Introduction and family Orthotrichaceae. Tropical Bryology. 11: 151-160.
  • Ramsay, H.P. & J.R. Spence. 1996. Chromosome data on Australasian Bryaceae. J. Hattori Bot. Lab. 80: 251-270.
  • Ramsay, H.P.1997. Cytotaxonomic studies Polytrichales from Australia, New Zealand, Papua New Guinea and Vanuatu. J.Hattori Bot. Lab. 82: 213-226.
  • Ramsay, H.P. 2006. Chromosome numbers in some species of Dicranoloma from Australia, New Zealand and Papua New Guinea. Telopea 11(3), 308-313.
  • Ramsay, H.P. 2007. Chromosome studies on some mosses from Papua New Guinea. Tropical Bryology.

FLORA of Australia – manuscripts completed and in press at ABRS

  • Ramsay, H.P., D.H. Vitt & J. Lewinsky-Hapaasori The family Orthotrichaceae.
  • Ramsay, H.P. & J.R. Spence. The family Bryaceae
  • Ramsay, H.P. Tan, B.C. & W.B. Schofield. The family Sematophyllaceae [the revision has been published ready for use.]
  • Ramsay, H.P.-Family Leptostomataceae.
  • Ramsay, H.P. with B. Allen - Family Dicnemonaceae.
  • Ramsay, H.P. Introduction to the Bryophytes (Mosses) Vol 51.
Helen Ramsay