|
1 |
Xanthones present |
2 |
| 1a |
Atranorin only present; spores 88-104 µm long; Thailand |
P. phukaensis Jariangprasert |
| 1b |
Xanthones absent |
38 |
|
2 |
Lichexanthone or derivatives present |
3 |
| 2a |
Norlichexanthone derivatives present [including thiophaninic acid] |
29 |
|
3 |
Lichexanthone present |
4 |
| 3a |
Chlorinated lichexanthone derivatives present |
11 |
|
4 |
Lichexanthone only present; spores 80-120 µm long; N.Am, Central Afr. |
P. valliculata Dibben |
| 4a |
Lichexanthone and stictic acid ± depsides or 2-chlorolichexanthone present |
5 |
|
5 |
Lichexanthone and 2-chlorolichexanthone present |
6 |
| 5a |
Lichexanthone and stictic acid ± depsides present |
7 |
|
6 |
With additional 2-O-methylsuperlatolic and 2-O-methylhyperlatolic acids; spores 90-115 µm long; Aus, S.Am, Cuba, PNG |
P. depressa (Fée) Müll. Arg.  |
| 6a |
With additional stictic acid; spores 85-120 µm long; Aus. |
P. atromaculata A.W. Archer & Elix |
|
7 |
Lichexanthone, constictic and stictic acids present; spores 75-130 µm long; S.Am |
P. rhodiza Nyl. |
| 7a |
Lichexanthone, stictic acid and depsides present |
8 |
|
8 |
Confluentic acid present |
9 |
| 8a |
Stenosporic acid derivatives present |
10 |
|
9 |
Spores 2-4/ascus, 80-176 µm long; Thailand |
P. inthanonensis Jariangprasert |
| 9a |
Spores 4-6-8/ascus, 87-110 µm long; S.Am. |
P. gracilis var. heteromera Müll. Arg. |
|
10 |
2'-O-Methylstenosporic acid present; spores 90-110 µm long; Thailand |
P. elixii Jariangprasert |
| 10a |
2,2'-Di-O-methylstenosporic acid present; spores (3-)4/ascus 76-92 µm long; Thailand |
P. alboaspera var. tetraspora Jariangprasert  |
|
11(3a) |
2-Chlorolichexanthone, stictic, 2-O-methylisohyperlatolic and 2-O-methylsuperlatolic acids present; spores 105-125 µm long; Aus. |
P. follmanniana A.W. Archer & Elix |
| |
2-Chlorolichexanthone and 2-O-methylsuperlatolic acid present; spores 100-120 µm long P438 |
var. nov. 'P. follmanniana var. toonumbarensis' |
| 11a |
4.5-Dichlorolichexanthone or 2,4,5-trichlorolichexanthone present |
12 |
|
12 |
4,5-Dichlorolichexanthone present |
13 |
| 12a |
2,4,5-Trichlorolichexanthone present |
26 |
|
13 |
4,5-Dichlorolichexanthone only present; spores 75-110 µm long; ostioles inconspicuous, pale brown, translucent; S.Am., Aus |
P. platystoma Malme |
| 13a |
4,5-Dichlorolichexanthone and depsidones and/or depsides present |
14 |
|
14 |
4,5-Dichlorolichexanthone and norstictic acid present; spores 80-110 µm long; S.Am; [H-NYL 3702 pm] |
P. 'rhodiza' Nyl.  |
| 14a |
4,5-Dichlorolichexanthone and stictic acid or depsides present |
15 |
|
15 |
4,5-Dichlorolichexanthone and depsides present |
16 |
| 15a |
4,5-Dichlorolichexanthone and stictic acid ± depsides present |
18 |
|
16 |
4,5-Dichlorolichexanthone and 2,2'-di-O-methylstenosporic acid present; spores 2-3(-4)/ascus 84-142 µm long; Thailand |
P. subplanaica var. tetraspora Jariangprasert & A.W.Archer |
| 16a |
4,5-Dichlorolichexanthone and 2'-O-methylstenosporic acid; spores 80-100 µm; Thailand |
P. kansrii Jariangprasert |
| 16b |
4,5-dichlorolichexanthone and 4-O-demethylplanaic acid present; spores 100-156 µm long; Thailand |
P. khuntanensis Jarinagprasert |
| 16c |
4,5-Dichlorolichexanthone and perlatolic acid derivatives present |
17 |
17
|
4,5-Dichlorolichexanthone and planaic acid present; spores (1-)2-3/ascus 90-174 µm long; Thailand |
P. siamensis Jariangprasert
|
| |
4,5-Dichlorolichexanthone, planaic and 2-O-methylperlatolic acids present; spores 4/ascus, 95-125 µm long; Aus |
P. doradorensis Elix & A.W. Archer |
| 17a |
4,5-Dichlorolichexanthone and 2-O-methylperlatolic acid present; spores 80-94 µm long; Aus, S.Am |
P. malmeii Elix & A.W. Archer  |
|
Spores 64-78 µm long; Aus. |
P. wilsoneana A.W. Archer & Elix  |
| 17b |
4,5-Dichlorolichexanthone and 2'-O-methylanziaic acid present; spores 84-146 µm long; Thailand |
P. phusoidaoensis Jariangprasert.  |
| 17c |
4,5-Dichlorolichexanthone and 2-O-methylisohyperlatolic; spores 105-115 µm; PNG |
P. streimannii Elix & A.W. Archer  |
| 18 |
4,5-Dichlorolichexanthone and stictic acid only present |
19 |
| 18a |
4,5-Dichlorolichexanthone, stictic acid and depsides present |
21 |
| 19 |
Thallus isidiate; spores 90-160 µm long; Eur. |
P. coronata (Ach.) Th.Fr.  |
| 19a |
Thallus not isidiate |
20 |
| 20 |
Spores 2-3(-4)/ascus; 75-100 µm long; Tahiti; verrucae confluent |
P. hypochrysea Vain. |
|
Spores 2-4/ascus, 51-140 µm long; pantemperate, N. hemisphere; Hawaii; verrucae flattened |
P. leucostoma (Bernh.) Massal.  |
| 20a |
Spores 60-90 µm; Eur. long; verrucae contricted at base |
P. caesioumbrina Eitner |
| 20b |
Spores 150-185 µm long, (2-)4(-5) per ascus; Japan |
P. glauca Zahlbr. |
| 21 |
4,5-Dichlorolichexanthone, stictic acid and perlatolic or stenosporic acid derivatives present |
22 |
| 21a |
4,5-Dichlorolichexanthone, stictic acid and 2-O-methylconfluentic acid present; spores (3-)4/ascus, 85-125 µm long; PNG |
P. bogia A.W. Archer & Elix |
| 21b |
4,5-Dichlorolichexanthone, stictic acid and confluentic acid present; spores 75-90 µm long; India |
P. cinchonae Müll. Arg. |
| 22 |
4,5-Dichlorolichexanthone, stictic acid and perlatolic acid derivatives present |
23 |
| 22a |
4,5-Dichlorolichexanthone, stictic acid and 2'-O-methylstenosporic acid present; spores 3-4/ascus, 80-108 µm long |
P. thailandica Jariangprasert |
| 23 |
4,5-Dichlorolichexanthone, stictic acid and planaic acid present; spores 82-100 µm long; S.Am. |
P. bonariensis Malme |
| 23a |
4,5-Dichlorolichexanthone, stictic acid and 2-O-methylperlatolic or 2'-O-methylperlatolic acidpresent |
24 |
|
24 |
Containing 2'-O-methylperlatolic acid; spores 90-150 µm long; Aus |
P. hermaka A.W. Archer |
| 24a |
Containing 2-O-methylperlatolic acid |
25 |
| 25 |
Spores (2-)4/ascus, 163-200 µm long; PNG, Philippines |
P. copelandii Vain. |
| 25a |
Spores 4/ascus, 100-120 µm long; PNG |
P. laeana A.W. Archer & Elix |
| 26(12a) |
2,4,5-Trichlorolichexanthone, 2-O-methylperlatolic and stictic acids; spores (2-)3(-4)/ascus, 70-100 µm long; Aus |
P. tjaetabensis A.W. Archer & Elix |
| |
2,4,5-Trichlorolichexanthone, 2'-O-methylperlatolic and stictic acids; spores 100-130 µm long |
P 360 Pertusaria sp. |
| 26a |
2,4,5-Trichlorolichexanthone and 2,5-dichlorolichexanthone present |
27 |
| 27 |
Additional 2-O-methylperlatolic acid present; spores 70-80 µm; Aus |
P. aquilonia A.W. Archer & Elix |
| 27a |
Additional stictic acid present |
28 |
| 28 |
Additional confluentic acid present; spores (3-)4/ascus, 64-75 µm long; Aus |
P. ewersii A.W. Archer & Elix  |
| 28a |
Depsides absent; spores 95-125 µm long; Sri Lanka, Java, PNG, Aus. |
P. ceylonica Müll. Arg.  |
| 29(2a) |
Arthothelin and derivatives present |
30 |
| 29a |
Thiophaninic acid present |
31 |
30
|
6-O-Methylarthothelin, arthothelin and stictic acid present; spores 117-125 µm long circumtropical? |
P. quasiae (Fée) Nyl. 
|
| 30a |
6-O-Methylarthothelin and arthothelin present; spores 45-62 µm long; PNG |
P. inconspicua A.W. Archer & Elix  |
| 31 |
Stictic or norstictic acid present, or stictic acid absent |
32 |
| 31a |
Depsides or anthraquinone present |
37 |
| 32 |
Thiophaninic and norstictic acids present |
33
|
| 32a |
Stictic acid present or absent |
34 |
| 33 |
Spores 68-110 µm long; N.Am |
P. wulfenoides de Lesd.
|
| 33a |
Spores 85-120 µm long; Socotra |
P. socotrana Müll. Arg. |
| 34 |
Lacking stictic acid |
35 |
| 34a |
With stictic acid |
36 |
| 35 |
2'-O-Methylperlatolic acid present; spores 4/ascus, 95-120 µm long: PNG |
P. rechingeri Zahlbr. |
| 35a |
Glomelliferic acid present; spores (2-)3/ascus, 80-120 µm long; Aus |
P. glomellerifica Elix & A.W. Archer  |
| 35b |
2-O-Methylperlatolic acid or 2,2'-di-O-methylstenosporic acid or confluentic acid present; spores 70-110 µm long; ostioles black, conspicuous; Aus |
P. trimera (Müll. Arg.) A.W. Archer
|
| 35c |
Thiophaninic acid only present; spores 4/ascus, 70-80 µm; New Caledonia, Thailand |
P. endochroma Müll. Arg. |
| 36 |
Spores (3-)4/ascus, 75-90 µm long; Aus, China |
P. abberans Müll. Arg. |
| 36a |
Spores 4/ascus, 100-130 µm long; Japan |
P. boninensis Shibuichi |
| 37(31a) |
Unknown anthraquinone present [K+violet]; spores 4/ascus, 100-160 µm long; Japan also saxicolous |
P. kashiwadanii Shibuichi |
| 37a |
4-O-Methylisocryptochloropheaic acid present; spores 4/ascus, 95-125(-145) µm long |
P. paradoxica A.W. Archer & Elix var. tetraspora A.W. Archer & Elix |
| 38(1a) |
Stictic or norstictic acid present |
39
|
| 38a |
Stictic and norstictic acid absent |
44 |
| 39 |
Stictic acid present |
40 |
| 39a |
Norstictic acid present |
43 |
| 40 |
Spores 100-150 µm long, walls rough; Japan |
P. radiata Oshio |
| |
Spores 140-160 µm, walls smooth; West Indies |
P. dussii Vain.  |
| 40a |
Stictic acid and perlatolic or stenosporic acid derivatives |
41 |
| 41 |
Perlatolic acid derivatives present |
42 |
| 41a |
Methyl 2'-O-methylstenosporate present; spores 2-3/ascus 118-150 µm long; Thailand |
P. methylstenosporica Jariangprasert |
| 42 |
2-O-Methylperlatolic acid present; spores 2-(3-4)/ascus, 84-130 µm long; Thailand; Aus |
P. loeiensis Jariangprasert
|
| 42a |
2'-O-Methylperlatolic acid present; spores 115-150 µm long; PNG |
P. novaeguineae A.W. Archer & Elix |
| 43(39a) |
Spores 85-110 µm long; Hawaii [K++red] |
P. ruboreagens H. Magn. |
| 43a |
Spores 62-85 µm; ostioles pale; S.Am |
P. cryptocarpa Nyl. |
|
Spores 65-85 µm; ostioles black; PNG |
P. naduriensis A.W. Archer & Elix  |
| 44(38a) |
Lichen compounds present |
45
|
| 44a |
Lichen compounds absent |
50 |
| 45 |
2-O-Methylsuperlatolic, 2'-O-methylisohyperlatolic or protocetraric acid present |
46 |
| 45a |
2'-O-Methylperlatolic or confluentic acid present |
49 |
| 46 |
Protocetraric acid present; spores 16-22 µm long; S.Am |
P. huanicola Messuti & A.W. Archer
|
| 46a |
Perlatolic acid or its derivatives present |
47 |
| 47 |
Perlatolic acid and fumarprotocetraric acid present; spores 90-150 µm long; Japan |
P. nakamurae (Räs.) Dibben  |
| 47a |
Derivatives of perlatolic acid present |
48 |
48
|
2-O-Methylsuperlatolic acid present; spores 3-(4)/ascus, 80-100 µm long; PNG |
P. manamensis A.W. Archer & Elix
|
| 48a |
2'-O-methylisohyperlatolic acid present; spores 65-69 µm long; Mexico |
P. polystictoides Vain.
 |
| 49 |
Confluentic acid present; spores 2-3(-4)/ascus, 72-134 µm long; Thailand |
P. orientalis Jariangprasert |
| 49a |
2'-O-Methylperlatolic acid present; spores (2-)3/ascus, 140-168 µm long; China |
P. parapycnothelia Q. Ren & Z.T. Zhao |
| 50(44a) |
In S.Am; spores 80-100 µm long |
P. polita var. tetramera Müll. Arg.
|
| 50a |
In Australasia |
51 |
| 51 |
In PNG; spores 90-110 µm long; ostioles inconspicuous, translucent |
P. plethocarpa A.W. Archer, Elix & Streimann  |
| 51a |
In Norfolk Island; spores 95-115 µm long; ostioles black, lacking a hyaline margin |
P. nebulosa A.W. Archer |